Cyclanorbinae

Subfamily of turtles

Cyclanorbinae Temporal range: Late Cretaceous to recent, 70.6–0 Ma PreꞒ Ꞓ O S D C P T J K Pg N

Indian flapshell turtle Lissemys punctata
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Testudines
Suborder:	Cryptodira
Family:	Trionychidae
Subfamily:	Cyclanorbinae Lydekker, 1889

Cyclanorbinae, also known commonly as the flapshell turtles, is a subfamily of softshell turtles in the family Trionychidae. The subfamily is native to Africa and Asia.

Taxonomy

Morphological evidence supports Cyclanorbinae being the sister group to the Plastomeninae, an extinct subfamily of softshell turtles that inhabited North America from the Cretaceous to the Eocene.^[1]^[2]

The subfamily Cyclanorbinae contains the following three extant genera.^[3]

One extinct prehistoric genus is also known from fossil remains: Nemegtemys Danilov et al., 2014, from the Cretaceous of Mongolia.^[4]

Species in the genera Cyclanorbis and Cycloderma are found in Africa; species in the genus Lissemys are found in Asia.^[5]

^ Joyce, Walter G.; Revan, Ariel; Lyson, Tyler R.; Danilov, Igor G. (October 2009). "Two New Plastomenine Softshell Turtles from the Paleocene of Montana and Wyoming". Bulletin of the Peabody Museum of Natural History. 50 (2): 307–325. doi:10.3374/014.050.0202. ISSN 0079-032X. S2CID 85505337.
^ Jasinski, Steven E.; Heckert, Andrew B.; Sailar, Ciara; Lichtig, Asher J.; Lucas, Spencer G.; Dodson, Peter (2022-07-01). "A softshell turtle (Testudines: Trionychidae: Plastomeninae) from the uppermost Cretaceous (Maastrichtian) Hell Creek Formation, North Dakota, USA, with implications for the evolutionary relationships of plastomenines and other trionychids". Cretaceous Research. 135: 105172. Bibcode:2022CrRes.13505172J. doi:10.1016/j.cretres.2022.105172. ISSN 0195-6671. S2CID 246803273.
^ Turtle Taxonomy Working Group (Rhodin, Anders G.J.; Parham, James F.; van Dijk, Peter Paul; Iverson, John B.) (2009)."Turtles of the World: Annotated Checklist of Taxonomy and Synonymy, 2009 Update". Archived 2016-04-06 at the Wayback Machine Chelonian Research Monographs (5): 000.39-000.84. (Cyclanorbinae, pp. 000.55-000.56).
^ "Fossilworks: Nemegtemys ". fossilworks.org. Retrieved 17 December 2021.
^ "Cyclanorbinae". The Reptile Database. www.reptile-database.org.

Cheloniidae	†Allopleuron Caretta †Carolinochelys Chelonia †Eochelone Eretmochelys †Gigantatypus †Glarichelys †Itilochelys Lepidochelys †Mexichelys †Miocaretta Natator †Pacifichelys †Syllomus †Tasbacka
Dermochelyidae	†Arabemys †Corsochelys †Cosmochelys Dermochelys †Eosphargis †Mesodermochelys †Psephophorus
	†Euclastes †Peritresius †Procolpochelys †Protosphargis †Puppigerus

Dermatemydidae	Dermatemys
Kinosternidae	Claudius †Hoplochelys Kinosternon Staurotypus Sternotherus

Emydidae	†Acherontemys Chrysemys Clemmys Deirochelys Emys Actinemys Emydoidea Glyptemys Graptemys Malaclemys Pseudemys Terrapene Trachemys †Wilburemys
Geoemydidae	Batagur †Banhxeochelys Cuora Cyclemys Geoclemys Geoemyda Hardella Heosemys Leucocephalon Malayemys Mauremys Melanochelys Morenia Notochelys Orlitia Pangshura Rhinoclemmys Sacalia Siebenrockiella Vijayachelys
Platysternidae	Platysternon
Testudinidae	Aldabrachelys Astrochelys Centrochelys Chelonoidis Chersina Cylindraspis †Cymatholcus †Floridemys Geochelone Gopherus †Hadrianus †Hesperotestudo Homopus Indotestudo Kinixys Malacochersus Manouria †Megalochelys †Oligopherus Psammobates Pyxis †Solitudo Stigmochelys †Stylemys Testudo

Carettochelyidae	†Allaeochelys †Anosteira Carettochelys
Trionychidae	Amyda Apalone †Axestemys Chitra Cyclanorbis Cycloderma †Drazinderetes Dogania †Gilmoremys †Hutchemys †Khunnuchelys Lissemys Nilssonia Palea †Palaeoamyda Pelochelys Pelodiscus Rafetus Trionyx
	†Basilochelys †Sinaspideretes

Chelydridae	Chelydra †Chelydrops †Chelydropsis †Emarginachelys †Macrocephalochelys Macrochelys †Planiplastron †Protochelydra
†Nanhsiungchelyidae	†Anomalochelys †Basilemys †Jiangxichelys
†Protostegidae	†Alienochelys †Archelon †Atlantochelys †Bouliachelys †Calcarichelys †Cratochelone †Desmatochelys †Iserosaurus †Notochelone †Ocepechelon †Pneumatoarthrus †Protostega †Rhinochelys †Santanachelys †Terlinguachelys
	†Adocus †Argillochelys †Bashuchelys †Ctenochelys †Prionochelys †Toxochelys

†Araripemydidae	†Araripemys
†Bothremydidae	†Araiochelys †Arenila †Azabbaremys †Bothremys †Cearachelys †Chedighaii †Chupacabrachelys †Eotaphrosphys †Foxemys †Galianemys †Ilatardia †Inaechelys †Itapecuruemys †Jainemys †Kinkonychelys †Kurmademys †Labrostochelys †Nigeremys †Phosphatochelys †Polysternon †Puentemys †Rosasia †Rhothonemys †Sankuchemys †Taphrosphys †Ummulisani †Zolhafah
Chelidae	Acanthochelys Chelodina Chelus Elseya Elusor Emydura Hydromedusa †Lomalatachelys Mesoclemmys Myuchelys Phrynops Platemys †Prochelidella Pseudemydura Rheodytes Rhinemys †Yaminuechelys
Pelomedusidae	Pelomedusa Pelusios
Podocnemididae	†Albertwoodemys †Bauruemys †Brontochelys †Caninemys †Carbonemys †Cerrejonemys †Cordichelys Erymnochelys †Lapparentemys †Latentemys Peltocephalus Podocnemis †Stupendemys
†Sahonachelyidae	†Sahonachelys †Sokatra

	†Caribemys †Caririemys †Tacuarembemys

Phylogenetic arrangement of turtles based on Turtles of the World 2017 Update: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status. † = extinct.